Marshall, T. C.; Coltman, D. W.; Pemberton, J. M.; Slate, J.; Spalton, J. A.; Guinness, F. E.; Smith, J. A.; Pilkington, J. G.; Clutton-Brock, T. H. (2002). Estimating the prevalence of inbreeding from incomplete pedigrees. Proceedings of the Royal Society B: Biological Sciences, 269(1500), 1533–1539.
doi:10.1098/rspb.2002.2035
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